Specimens have been collected on the ground in spruce and lodgepole pine forests in the western mountains and in conifer forests of many kinds in the east. Occasional specimens ewre collected in oak, poplar, or birch forests and in salt marshes, sphagnum bogs, and beach litter along the shores of seas and lakes. (B163); primarily boring-old forest but also decid forests, salt marshes, bogs, and in litter on lake and sea shores (R197);